วันพฤหัสบดีที่ 17 กันยายน พ.ศ. 2569

Laos limestone forest

 Khammouane 

ແຂວງ ຄໍາມ່ວນ


Khammouane province (Khammouan) (Lao: ຄໍາມ່ວນ, pronounced [kʰám mūan]) is a province in southern Laos. Its capital is Thakhek. Khammouane province covers an area of 16,315 square kilometres (6,299 sq mi), most of which is forested mountainous terrain. The forests of the Nakai-Nam Theun National Park contains streams that feed Mekong tributaries and form the catchment area for Nam Theun 2, the largest hydropower project in Laos. The Xe Bang Fai River, Nam Hinboun River, and the Nam Theun are some of the rivers of the province.

Part of the aborted Thakhek-Tan Ap railway would have crossed the province to connect with the North-South Railway at Tân Ấp Railway StationQuảng Bình provinceVietnam through Mụ Giạ Pass. Khammouane's human development index is 0.602 which is less than the Laos average of 0.620.

Khammouane province covers an area of 16,315 square kilometres (6,299 sq mi)and is mostly of forested mountainous terrain.The province is bordered by Bolikhamsai province to the north and northwest, Vietnam to the east, Savannakhet province to the south, and Thailand to the west.[ Streams flow through the province to join the Mekong River. Some rivers originating in the mountains of the province are the Xebangfay River (239 kilometres (149 mi)long), Nam HinbounNam Theun, and Nam Ngum River. Between the Mekong and the Annamite Range, the Khammouane Plateau features gorges, grottoes, jungles, limestone hills, and rivers.

The Rock Viewpoint at Phou Pha Marn




The Rock Viewpoint opens the doors to the Phou Hin Boun National Protected Area, an uncharted world of ancient karstic forests, breath-taking views, and endemic wildlife in Central Laos. Immerse yourself in one of the largest limestone forest in all of Asia with a fascinating itinerary of zip lines, spider net and canopy walks. The viewpoint, known as “The Peak of the loop” is a must stop on the Thakek Loop that also acts as a tourist information centre for all visitors in Khammouane Province. Whether having a coffee at the Coffee Bar, delicious meal at the Khob Chai Deu restaurant or an incredible adventure is what you seek, the stunning views are assured!

Discover the Limestone Forest of Phou Pha Marn and experience the Highlights of The Rock for yourself! This experience is perfect for those who want to enjoy the highlights of the Protected Area within a short time. Accessible for almost everybody, it is recommended for families and friends. Start the tour with an epic zip line, flying across the valley of this stunning landscape, to the thrilling circuit of via ferrata, suspended bridges, ziplines and spider net. The trip ends with an incredible fly across the rock formations on the final zipline, back to the Phou Pha Marn visitor centre where you can enjoy the views from the Khop Chai Der restaurant.

Gesneriaceae
Laos is home to a rich diversity of Gesneriaceae (the African violet family), with numerous newly discovered endemic species and recent botanical records identified in limestone karsts and evergreen forests
Newly Described Species
Recent scientific expeditions and studies have documented several species unique to Laos:
  • Didymocarpus species: Didymocarpus angustiflorusD. bolavenensisD. laoticus, and D. trilobus.
  • Paraboea species: Paraboea planiflora and Paraboea khotamiae, discovered in the limestone karsts of central Laos.
  • Other novelties: Hemiboea olivifoliaChayamaritia banksiae (endemic to Khammouane Province), and Lysionotus calcicola found across northern and central provinces. 
Recent Floral Records
Botanists have also recorded species previously thought to be restricted to neighboring countries or new to the Pha Khao Lao database: 
  • Damrongia trisepala and Didymocarpus formosus (previously considered endemic to Thailand)
  • Middletonia reticulata
  • Aeschynanthus bracteatus and A. micranthus
  • Epithema ceylanicumLysionotus serratusMicrochirita luteolaParaboea swinhoeiPetrocosmea kerrii, and Pseudochirita guangxiensis 

Several species of the genus Paraboea (family Gesneriaceae) are found in the limestone karsts of Khammouane Province, central Laos.
Notably, these include:
  • Paraboea khotamiae: A micro-endemic species discovered and described in 2021. It is known to grow specifically on shaded limestone cliffs in the Gnommalat District of Khammouane. It features a white corolla and non-twisted seed capsules.
  • Paraboea mahaxayana: A species named after the Mahaxay region in Khammouane, where it grows in the fissures of large limestone rocks

Paraboea khotamiae  ດອກລະຄັງໂຄຕະມີ 


Recently Described Species in Khammouane
  • Microchirita argostemmifloraMicrochirita flavofusca, and Microchirita magniflora: Three distinct new species of Microchirita were described from the Khammouane Karst in 2024 as part of ongoing botanical explorations of Lao Gesneriaceae. 
Microchirita argostemmiflora

Microchirita magniflora:
Microchirita flavofusca


  • Lysionotus calcicola: A deciduous, shrubby species adapted to severe limestone cliff environments, found distributed across Luangprabang, Vientiane, and the Khammouane provinces.

Lysionotus
 D.Don (1822, p. 85), belonging to the family Gesneriaceae (subfamily Didymocarpoideae), consists of 35 species distributed from northern India and Nepal eastwards through Bangladesh, Bhutan, Laos, Myanmar, Thailand, northern Vietnam and southern China, Taiwan to southern Japan (Möller et al. 2017, GRC 2024 continuously updated2024 continuously updated). In Laos, only two species, L. levipes (C.B.Clarke) B.L.Burtt and L. serratus D.Don have been found (Newman et al. 2017 onwards2017 onwards, Souvannakhoummane et al. 2021). The genus is characterized by its sub-shrubby habit, opposite or whorled leaves, tubular funnel-shaped corolla with an inflated upper half, two stamens, coherent anthers with or without connective appendages, and seeds with an appendage at each end (Weber et al. 2020, Akhil et al. 2021).

Khammouane Province is one of the richest areas with karst topography in Laos. Clusters of limestone hills topped with limestone caves make the province famous, such as The Rock Viewpoint at Phou Pha Marn, Konglor Cave in Phu Hin Poun National Protected Area and Xe Bang Fai Cave in Hin Nam No National Park, and others. Supported by this topography, the Lao limestone karsts provide specific habitats for various plant species endemic to this area, such as Gesneriaceae: Didymocarpus middletonii Souvann., Soulad. & Tagane (Souvannakhoummane et al. 2018), Paraboea khotamiae Phonep. & Souvann. (Phonepaseuth et al. 2021), Poaceae: Laobambos calcareus Haev., Lamxay & D.Z.Li. (Haevermans et al. 2020), Polypodiaceae: Drynaria deuansavanhii Vongthavone & Tagane, Pteridaceae: Hemionitis khammouanensisVongthavone & Phonep. (Vongthavone et al. 2025), and Vitaceae: Tetrastigma fruticosumSoulad., Phonep. & Tagane (Tagane et al. 2025).

Recently, an unknown species of Gesneriaceae was discovered and collected during botanical surveys in the limestone karst of the Rock Viewpoint at Phou Pha Marn, Khammouane Province. From our morphological observations, the plant was assignable to the genus Lysionotus D.Don based on seeds with an appendage at each end (Weber et al. 2020, Akhil et al. 2021). The new species is similar to Lysionotus chingii Chun ex W.T.Wang by being a subshrub with large calyx and corolla, and these two species are compared in details. In addition, the same species was collected also on the limestone of Pha Yoak Limestone Forest Reserve Area, Luangprabang Province, northern Laos and Vangvieng District, Vientiane Province, Mahaxay District, Khammouane Province, central Laos. After careful examination of these specimens, morphological comparison with closely related species, and consultation of the relevant taxonomic literature, it was concluded to be an undescribed species. In addition to the morphological examination, DNA sequences and phylogenetic analysis are extremely helpful for delimiting species (Hebert and Gregory 2005, Dick and Webb 2012). Here, we sequenced a DNA barcode region, the nuclear ribosomal internal transcribed spacer (ITS) to compare the new species with related taxa. As a result, we describe it as new to science herein, along with the line drawing, information on its distribution, ecology, phenology, vernacular name, and provisional conservation status.

  • Didymocarpus middletonii: Recorded from the limestone karsts within the region's protected areas. 


THAM XANG CAVE (ELEPHANT CAVE)

Tham Xang Cave, commonly known as Elephant Cave, is one of the most interesting cultural and natural attractions in Vang Vieng. Located approximately 7 kilometers north of Vang Vieng town, near Tham Norn Cave and Tham NangFa (Angel Cave), this cave is famous for its unique limestone formation that resembles the head of an elephant, which gave the cave its name. Although smaller than some of Vang Vieng's larger cave systems, Tham Xang is valued for its religious significance, historical importance, and easy accessibility.


The cave is situated at the base of a dramatic limestone mountain surrounded by lush greenery and rural countryside. Visitors can reach the cave through a short and easy walk, making it suitable for travelers of all ages. The entrance opens into a cool chamber where the cave's most famous feature—a naturally formed limestone rock resembling an elephant—can be found. Local people have long regarded this formation as sacred, and it has become an important symbol of the cave.





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Laos limestone forest

  Khammouane   ແຂວງ ຄໍາມ່ວນ Khammouane province (Khammouan) ( Lao : ຄໍາມ່ວນ, pronounced [kʰám mūan] ) is a province in southern Laos. Its ca...